This is the result I get with a game base character using the Auto setup for 4.25 (Look at the nose and ears, you might need to view full size image), this happens with CC3 and CC3+ base characters and with normal export and InstaLOD.
I never had this issue before until I upgraded to 3.3
I get the same result without using CC setup and manually create the materials.
The mesh is around 17K triangles.

Any help is much appreciated, thanks.